Center Joe Berger (concussion) and free safety Harrison Smith (ankle) have been ruled out for Sunday’s game at Jacksonville, but punt returner Marcus Sherels (rib) is questionable after missing the past two games, one of which turned in the wrong direction on Adam Thielen’s muffed punt inside the Vikings’ 10-yard line.

Also ruled out is defensive tackle Sharrif Floyd, who has been sidelined since the season opener because of a knee injury.

Also listed as questionable is backup linebacker/special teamer Edmond Robinson.

Second-year pro Anthony Harris is likely to step in for Smith. Nick Easton will make his second straight start for Berger.

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Harris gets additional work during Vikings practice without Smith

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Sherels, Robinson downgraded to out for Vikings in Jacksonville