Florida and coach Jim McElwain agreed to part ways a day after a third consecutive loss and nearly a week after he said his players and their families had received death threats.
Athletic director Scott Stricklin made the announcement Sunday and named defensive coordinator Randy Shannon the team's interim coach for the final four games.
The parties are negotiating McElwain's $12.5 million buyout.
Florida (3-4, 3-3 Southeastern Conference), which is still paying former coach Will Muschamp, would like to significantly reduce McElwain's sum and could use his actions over the last week as leverage.
Saturday's 42-7 loss to Georgia dropped McElwain's record to 22-12 with the Gators, including 4-9 against ranked teams.
AP Top 25 poll
• Iowa State made the biggest jump in the Associated Press Top 25 poll after its victory over TCU, climbing 11 spots to No. 14. The Cyclones have their highest ranking since being No. 9 on Oct. 13, 2002.
• Every team in the top 10 except No. 1 Alabama changed positions. In the Big Ten, Ohio State moved from sixth to third, Wisconsin from fifth to fourth and Penn State from second to seventh.
• Georgia is the new No. 2 team, and Notre Dame rounded out the top five.